Fula, Wolof or Temne? Upper Guinean AncestryDNA results 2013-2018

This blog post will feature a summary of my Upper Guinean AncestryDNA survey findings. A fitting conclusion of my African AncestryDNA research as I am myself of Cape Verdean descent. And therefore this particular section was of paramount significance to understand my own African Roots! These results were obtained during 2013-2018 but I had not fully processed all the data up till now. Relatively few African customers of Ancestry are hailing from this particular area. Which is why it was difficult to gather a sufficient sample size. But eventually I did succeed. Also through the valuable help of several friends!1 Follow the link below for detailed analysis & screenshots:

Table 1 (click to enlarge)

Stats, SEN, n=34

“Senegal” is primary for most countries throughout the wider Upper Guinea area. Usually with “Mali” as secondary region. This goes even for the northern part of Sierra Leone. But this country shows greater variation. With “Ivory Coast/Ghana” also being a prominent component overall. An almost equal “Mali” group average (38-39%) was obtained for 6 samples from Mali when compared with 3 Gur/Senoufo speaking samples from Burkina Faso, Ivory Coast & Ghana.

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Table 2 (click to enlarge)

Stats Fula

A clearly detectable Upper Guinean component is mirrored among the Fula from across Upper Guinea and the Hausa-Fulani. Indicating their shared Upper Guinean origins by way of eastwards moving Fula migrations. However due to their partial Nigerian Hausa lineage the Hausa-Fulani results can still be quite easily distinguished through their primary “Nigeria” scores.

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My final survey group now consists of 122 AncestryDNA results from Senegal, Gambia, Guiné Bissau, Guinea, Sierra Leone, Mali and Mauritania. Although still limited this data-set already provides a rather robust basis. All the more given that my combined survey group (n=122) contains a number of people which is almost three times greater than Ancestry’s Upper Guinean Reference Panel at that time (n=44; 28 samples being used for “Senegal” + 16 samples for “Mali”).

And also crucially I have managed to gather plausible ethnic details for many of my survey participants. Enabling me to compile a separate Fula survey group (n=46) which is quite extra-ordinary as it includes Fula people from a wide range of countries (see Table 2). While usually in published studies only Fula samples from one particular area are being covered (often from the central/eastern Sahel and not from Upper Guinea).

To a lesser extent I also uncovered more specific ethnic backgrounds among my Sierra Leonean and Senegambian survey groups. Allowing for a finer detailed ethnic analysis of Upper Guinean genetics. Which is why I think these “old” results may still be useful and are not obsolete yet. Even when I did discontinue this survey after Ancestry’s update in September 2018. Because in my opinion Ancestry’s new version of their ethnicity estimates regrettably has been a downgrade in regards to their African breakdown, overall speaking (see also this blogseries).

Five main implications for Afro-Diasporans can be singled out. These are discussed in greater detail on the main page. In this blog post I will mostly elaborate on the question if it is possible to distinguish Upper Guinean DNA. Furthermore I will briefly touch upon 23andme’s new “Senegambian & Guinean” category and also how Ancestry might improve things in the near future. Especially in light of their upcoming update.

  1. “Senegal” + “Mali” combined is a solid indication of lineage across Upper Guinea (Senegambia, Guinea Bissau/Conakry, Sierra Leone, western Mali).
  2. “Mali” can also be predictive of DNA found in Burkina Faso, northern parts of Ivory Coast & Ghana.
  3. “Ivory Coast/Ghana” can also be predictive of Sierra Leonean DNA
  4. “Africa North” might also be inherited by way of Fula ancestors
  5. “South-Central Hunter-Gathers” can also be predictive of West African ancestors

In summary: Regional admixture DOES matter! Judge each case on its own merits. Combine insights from different fields to achieve complementarity!

Figure 1 (click to enlarge)

COMPILUG

The two first results illustrate how AncestryDNA’s 2013-2018 version was able to clearly distinguish between Fula & Wolof lineage (for Africans themselves!). The additional “Africa North” and “Middle East” scores making the difference. To a lesser degree also within Sierra Leone some ethnic differentiation (going by group averages) could be observed. Obviously there was greater individual variation though. And in no way was either “Senegal” or “Mali” an exclusive marker for any given ethnic group! For Afro-Diasporans follow-up research is therefore required (DNA matches, historical plausibility, genetic genealogy etc.).

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Igbo, Yoruba or Hausa-Fulani? Nigerian AncestryDNA results 2013-2018

Without wanting to rehash things this blog post and following ones will provide a quick recap of my final AncestryDNA survey findings for continental Africans. These results were obtained during 2013-2018 but I had not fully processed all the data up till now. Moving on now to Nigeria, with a special focus on how to distinguish Igbo, Yoruba and Hausa-Fulani lineage.

I first published my preliminary Nigerian survey findings on 22 September 2016 when I had only 15 Nigerian AncestryDNA results available for my analysis (see this overview). As my survey has been ongoing I have managed to collect a sample group which is now five times greater. Consisting of no less than 87 AncestryDNA results of Nigerian persons! Follow the link below for detailed analysis & new screenshots:

Table 1 (click to enlarge)

naija comparison

For all three listed ethnic groups “Nigeria” is the primary regional component. However more differentiation is detectable when zooming into secondary regions. In particular “Senegal” for the Hausa-Fulani clearly stands out when compared with the rest. Less clear-cut distinction between Igbo & Yoruba. However when taking into account relative proportional shares for “Benin/Togo” and “Cameroon/Congo” it is still already detectable.

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I discontinued this survey after Ancestry’s update in September 2018. Because in my opinion Ancestry’s new version of their ethnicity estimates regrettably has been a downgrade in regards to their African breakdown. In particular Ancestry’s update in 2018 has been disastrous for obtaining reasonable Nigerian DNA results. Generally speaking former “Nigeria” scores have sharply decreased and were replaced by inflated “Benin/Togo” and “Cameroon, Congo, Southern Bantu” scores. Just as a reminder this blog post is NOT dealing with those updated and usually rather misleading results! Instead read this blogseries.

My Nigerian AncestryDNA survey is actually the most extensive and oldest part of my African survey (2013-2018). Such results initially being very difficult to come by. However currently my sample size (n=87) is rather robust. Higher even than Ancestry’s own Nigerian sample size (n=67) during this period! And also crucially I have managed to gather plausible ethnic details for almost all of my Nigerian survey participants. Allowing for a finer detailed ethnic analysis of Nigerian genetics. Which is why I think these “old” results may still be useful and are not obsolete yet. Even when I did already establish in 2016 that “Nigeria” does not not cover the full extent of one’s Nigerian lineage.

I originally singled out three main implications/propositions for Afro-Diasporans. The first two ones have been discussed already in previous blogs. However not so the last one which I will revisit in this blog post. Furthermore I will briefly touch upon 23andme’s new “Nigerian” category and also how Ancestry might improve things in the near future.

  1. “Benin/Togo” also describes DNA from Nigeria (see this blog post as well)
  2. “Cameroon/Congo” can also be partially indicative of southeastern Nigerian lineage (usually to a minor degree though, see this blog post)
  3. Is it possible to determine the most likely ethnic source(s) of your Nigerian lineage?

Figure 2 (click to enlarge)

Compil NG 3x

All three results show a predominant “Nigeria” amount. Indicative of a high degree of shared origins for Nigerians, regardless of ethnic background. Then again there is a major distinction between Hausa-Fulani and southern Nigerian results because of in particular the additional “Senegal” score and absence of “Benin/Togo” & “Cameroon/Congo”. Overlap between Yoruba and Igbo results is much greater but still going by proportional shares for in particular “Cameroon/Congo” still some minor differentiation can be detected.

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23andme’s new African breakdown put to the test

My first DNA test ever was with 23andme. Nine years ago already! In January 2010 I was thrilled but soon afterwards also quite underwhelmed to receive my very basic admixture results. The only distinction being made back then was between African, Asian and European DNA. Native American DNA did not even have a separate category yet 🙂 As I am of Cape Verdean descent I was actually most anxious to have my Upper Guinean lineage confirmed. Instead my African score just pointed towards the entire continent! One of my immediate reactions at that time therefore was:

“I hope that one day 23andme’s Ancestry Reports will be helpful in finding out where to locate my ancestry regionally and not just on a continental scale.”

After a (very) long wait it seems that this day has finally arrived! Last month 23andme rolled out an updated version (3.0) of Ancestry Composition to all their customers. Regardless of when they originally took the test. This update has actually been on release since September 2018 for 23andme’s most recent customers. But to its credit 23andme also made this update available to its earliest customers, like myself. Over the years I have been through more than one update on 23andme already. But this is the first time I can say that finally a meaningful African breakdown is being provided! For more details see:

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Figure 1 (click to enlarge)

23andmecompil

Updated 23andme results from across the African continent. A small but representative sample. Highlighting how 23andme’s new African regions appear to be quite predictive, for native Africans themselves. Unrealistic expectations about “100% accuracy” as well as counter-productive obsessing about regional labeling should be avoided. Instead take note of how the expected regions (circled in red by myself) reach levels of over 70% reaching into 98%! Taking a macro-regional perspective (combining overlapping regions from within West Africa versus Central/Southern Africa versus Northeast Africa) these results are usually in line as well. Also the additional ancestral locations appearing below the regional scores are on point!

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I have always believed that the best way to find out about the predictive accuracy of any particular DNA test or update is to look at the results of people who actually know their (recent) origins. In order to improve correct interpretation I have therefore started a survey among African DNA testers (n=173). Using their group averages as some sort of rudimentary benchmarks so to speak. Similar to the survey I conducted among African AncestryDNA testers in previous years (see this page). Of course also some basic knowledge about DNA testing (in particular 23andme’s reference populations and methodology) as well as historical context will remain essential to really get the most out of your admixture results!1

Main topics if you continue reading:

  1. Survey findings for 173 African 23andme testers from 31 countries (incl. 25 Cape Verdeans)
  2. Maps showing the geographical distribution of the new African regions on 23andme (based on my survey findings)
  3. Implications for Afro-Diasporans
  4. Examples to illustrate how regional admixture DOES matter!

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DNA matches reported for 50 Cape Verdeans on AncestryDNA (part 1)

In this two-part blogseries I will analyze the DNA matches being reported by AncestryDNA for 50 of my Cape Verdean survey participants. A follow-up to my previous blog post about 100 Cape Verdean AncestryDNA results (see this link). Because I was kindly given access to their profiles I was able to use my scanning and filtering method of DNA matches in Excel (see this link). Aside from matches with mainland Africans I am also including matches with people of (presumably) fully Portuguese, Jewish, West Asian and South Asian descent.1 Below a statistical overview of my main findings. Going by group averages. For the individual results which do display greater variation follow this link:

Table 1 (click to enlarge)

DNA matches for 50 CV's

This table is based on group averages. Except for the columns mentioning the frequency of close and zero matches. So for example among my 50 survey participants only one single person received a close African match (>20cM). While two persons did not receive any African matches at all (excl. North Africa). But on average 5 African matches were reported of whom 4 were connected to the Upper Guinea area. (Senegal-Sierra Leone). The average admixture amounts are based on the recently updated Ethnicity Estimates on AncestryDNA. This update strongly reduced the trace regions. Especially for North African & West Asian DNA. For a previous version of this table see this link.

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Table 2 (click to enlarge)

African matches

The background column is mostly based on informed speculation (plausible surnames/regional admixture) but at times also confirmed by public family trees. The proportion of Upper Guinean related matches is 88% of all African matches (south of the Sahara). That proportion being equal to 227/257. Excluding North African matches from the total. The high number of Fula matches is quite striking. But this could very well reflect a greater popularity of DNA testing among Fula people when compared with people from for example Guiné Bissau who are greatly underrepresented in Ancestry’s customer database.

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This project was merely intended as an exploratory exercise. Of course my research findings have limitations in several regards. And therefore they should be interpreted carefully in order not to jump to premature or even misleading conclusions. Still I do believe they can reveal relevant tendencies in DNA matching for Cape Verdeans in general. These outcomes may also provide valuable insight into the various ancestral components found within Cape Verdean DNA. In particular when aiming for complementarity by also taking in to account admixture analysis, genealogy and relevant historical context.

Below an overview of the topics I will cover in this blog post:

  1. Considerations when dealing with DNA matches
  2. Upper Guinean matches: as expected African matches (south of Sahara) were overwhelmingly from Upper Guinea (Senegal-Sierra Leone): 88% of the total. In line with the 92% Upper Guinean admixture proportion  (“Senegal” + “Mali” / total African) I found for my survey group.
  3. North African matches: fairly consistent despite minimal shared DNA
  4. Other African matches: unexpected & uncommon. Higher odds of false positives but in some cases to be corroborated by additional clues, such as AncestryDNA’s ethnicity estimates?
  5. Methodology: how I filtered the African DNA matches as well as the decision rules I applied when determining a plausible background for each DNA match.

Part 2 of this blogseries will have the following topics:

  1. Portuguese matches: omnipresent and clearly most numerous as well as often hinting at relatively recent ancestral ties (1800’s-1900’s).
  2. Jewish matches: Sephardi matches more likely to be truly genealogical than Ashkenazi matches?
  3. West Asian matches: quite rare, possibly indicating that West Asian admixture among Cape Verdeans is generally indicative of actual North African or Sephardi lineage.
  4.  South Asian matches: also rare, but on a hit and miss basis still sometimes already seemingly validating trace amounts of South Asian admixture.
  5. Inter-island matching patterns: illustrated by the distribution of the shared DNA segments between myself and my 100 Cape Verdean survey participants.
  6. Methodology: how I filtered the non-African DNA matches as well as the decision rules I applied when determining a plausible background for each DNA match.

Dedicated to all my Cape Verdean primos and primas participating in this survey.And special dedication to my newly born nephew Max!

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100 Cape Verdean AncestryDNA results

In October 2015 I published my first preliminary survey findings based on 23 Cape Verdean AncestryDNA results (see this link). Right now, almost three years later, I have managed to collect a sample group which is four times greater. Consisting of no less than 100 AncestryDNA results of fully Cape Verdean-descended persons! Even though this quadrupled sample size is obviously still limited it will most likely provide a greater insight in the various ways how “Caboverdeanidade” can be described. Genetically speaking that is. And obviously when applying the regional AncestryDNA format, with all its enhanced features as well as its inherent shortcomings  😉

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capeverdednalogo2

Click on this banner to reach Cape Verde DNA, Inc: the biggest online community of Cape Verdean Genealogy & DNA enthusiasts! On Augustus 4 & 5 a pioneering Cape Verde DNA and Genealogy Conference will be held! See this link for more details.

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In this blog post I will discuss the main differences with my previous findings from 2015, which were focused on the African breakdown solely. And in addition I will also present some new statistics and background information on the European and other non-African origins of Cape Verdeans as reported by AncestryDNABelow an overview of all the topics I will cover:

  1. Background details of my 100 Cape Verdean survey participants
  2. To be Cape Verdean is to be mixed?
  3. Upper Guinean roots = “Senegal” + “Mali”
  4. Beyond Upper Guinea: valid outcomes or misreading by AncestryDNA?
  5. European breakdown reflecting mostly Portuguese ancestry?
  6. “Africa North”, “Middle East”, “European Jewish” and other minor regional scores
  7. Upcoming update of AncestryDNA’s Ethnicity Estimates

Follow these links for my complete survey data & research methodology:

Table 1 (click to enlarge)

cvstats

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Chart 1 (click to enlarge)

Primary regions

This frequency of regions being ranked #1 (regions with the highest amount in either the African or European breakdown) is perhaps the best indicator of the main ancestral components for my Cape Verdean survey group. However only in an extra pronounced degree. For more nuance see the group averages in the next sections.

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Screenshots of individual results (rightclick and open in new tab to enlarge; island origins shown below)

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More charts and analysis when you continue reading!

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Update: Afro-Diasporan AncestryDNA Survey (part 1)

In 2013 AncestryDNA updated their Ethnicity Estimates to include a detailed breakdown of West African DNA. Pioneering when compared with other DNA testing companies. Soon afterwards I started collecting AncestryDNA results in an online spreadsheet in order to conduct a survey of the African regional scores being reported by AncestryDNA. At first only for people of the Afro-Diaspora and later on also among Africans. My main research goal has always been to establish how much the AncestryDNA results on an aggregated group level can already (despite limitations of sample size and other shortcomings) be correlated with whatever is known about the documented regional African roots for each nationality. As well as to improve correct interpretation of personal results.

In May 2016 I published my first summary of my Afro-Diasporan survey findings based on 707 results for 7 nationalities (see this blog post). My survey has been ongoing ever since. Right now an update of AncestryDNA’s Ethnicity Estimates seems even more imminent than it was in 2016 (when it was canceled in the beta phase). So that’s why I will yet again provide a “final” overview of my survey findings 😉 . Mainly based on 1,264 results for people from 8 nationalities. Although the total number of results and nationalities in my survey is even greater.

A major addition is the inclusion of 45 Brazilian results. Their predominant Central African profiles (as measured by both “Southeastern Bantu” and “Cameroon/Congo”) are quite striking when compared with my other sample groups. This outcome reinforces how the African breakdown on AncestryDNA has been reasonably in alignment with historically documented origins of the Afro-Diaspora. Unlike any other DNA testing platform I’m aware of and therefore not to be lightly dismissed despite inherent imperfections.

In the second part of this blogseries I will also provide an overview of the non-African regions (Amerindian, Asian, Pacific etc.) being reported for Afro-Diasporans. As well as a more detailed analysis of their European breakdown.

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This frequency of regions being ranked #1 (regions with the highest amount in the African breakdown) is perhaps the best indicator of which distinct African lineages may have been preserved the most among my sample groups.”

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Chart 1 (click to enlarge)

Afro piechartsa

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AncestryDNA Results Across the Diaspora

In 2013 AncestryDNA updated their Ethnicity Estimates to include a very detailed breakdown of West African ancestry (see this article). Soon afterwards I started collecting AncestryDNA results in an online spreadsheet in order to conduct a survey of the African regions being reported by AncestryDNA, among both African Americans as well as other Afro-descended nationalities. Attempting to establish how much the AncestryDNA results on an aggregated group level can already (despite limitations of sample size) be correlated with whatever is known about the documented regional African roots for each nationality.

Rumour has it that AncestryDNA will shortly start rolling out a new update of their Ethnicity Estimates. So it seems the time is right to finalize my survey. The sample size for most groups appears to be suffciently robust now to allow a meaningful intercomparison. In the AncestryDNA section of my blog (see the menubar) you can find a detailed summary of my survey findings based on 707 results for 7 nationalities:

Gathering all the results was a great learning experience. It has been a very satisfactory project! My survey report merely represents my personal attempt at identifying generalized, preliminary and indicative patterns on a group level inspite of individual variation. Everyone has a unique family tree of course first of all.

I would like to thank again all my survey participants for sharing their results with me. I am truly grateful for it!

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This frequency of regions being ranked #1 (regions with the highest amount in the African breakdown) is perhaps the best indicator of which distinct African lineages may have been preserved the most among my sample groups.”

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FREQ #1 regions

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Louisiana: most African diversity within the United States?

Louisiana Slave Database 1719-1820

SUMMARY

Number of slaves with origin specified 29,769
American born (“Creole”) 13,618 (45% of total)
African born 16,099 (54% of total)
African specified ethnically 8,994 (30% of total)

TOP 3 BREAKDOWN OF AFRICAN BORN SLAVES

Congo (Central Africa) 2,988 – 33% of African specified
Mandinga (Upper Guinea) 922 – 10% of African specified
Mina (Ghana, Togo, Benin) 628 – 7% of African specified
Source: Slavery and African Ethnicities in the Americas: Restoring the Links (Hall, 2005).

Congo Square

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Gwendolyn Midlo Hall’s remarkable relational database of Louisiana slaves and freedmen (1719-1820), which contains records of over 100,000 enslaved people, includes nearly 9,000 individual Africans by specific ethnicity. The database was first published in 2000 as a CD-ROM, and it is now available free on the Internet. The database is derived largely from notarial and ‘‘succession’’ (probate) records, mostly in French, in which slaves often self-identified their ‘‘nations’’ or ‘‘countries’’. There are 217 different ethnicities recorded, of which 96 have been identified, and a further 121 (comprising 152 individuals) which have not been identified. Of those identified, just 18 ethnicities account for over 96% of the Africans in the Louisiana records. In addition, within that set of 18 principal ‘‘nations,’’ the seven most common (in order, Kongo, Mandingo, Mina, Senegal/Wolof, Igbo, Bamana, Chamba), comprise over three-quarters of the sample. The single most numerous ‘‘nation’’ in the dataset was Africans from Kongo (nearly 3,000 individuals), the vast majority of whom arrived in the late eighteenth and early nineteenth centuries” (Chambers, 2008, pp.335-336).

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For a direct link to this extremely fascinating and online searchable database created by Gwendolyn Midlo Hall see below:

Louisiana Slave Database 1719-1820

A full overview of the 18 most frequently named African origins can be seen in the remaining part of this blogpost in which i will attempt to provide more details and context about Louisiana’s African diversity. Continue reading

Cape Verde Slave Census of 1856 (part 2)

Origins from across Upper Guinea, not just from Guinea Bissau

ic_image_ugc_new_jpg_1116049996

Map of Upper Guinea, western Mali should also be included for ancestral purposes

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Bissau, Cacheu, Cape Verde Slave Census of 1856

REGIONAL ORIGINS (combined)

Total ethnically specified: 1,615
Guinea Bissau’s Coastal Zone: 843 (52% of ethnically specified)
Upper Guinea Interior: 670  (42% of ethnically specified)
Senegal, Guinea & Sierra Leone: 102 (6% of ethnically specified)

TOP 3 ETHNIC ORIGINS (combined)

Mandinga (Upper Guinea) 262 – 16% of ethnically specified
Tilibonca (Upper Guinea) 229 – 14% of ethnically specified
Bijago (Guiné Bissau) 226 – 14% of ethnically specified
Source: Hawthorne (2003)

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In the first part of this blogpost i already discussed the main Guinean Bissau origins for Cape Verde according to its 1856 slave census, in this second part i will continue exploring origins outside of Guiné Bissau. When asked about their mainland African roots many Cape Verdeans might assume they only have ancestry coming from Guiné Bissau, this is however not completely true. It’s indeed correct that Guiné Bissau shares a very long and intimate history with Cape Verde. Both countries being ex-Portuguese colonies, united in their independence struggle during the 1970’s. Because of ever increasing English and French encroachment the formal Portuguese influence area within Upper Guinea during the 1600’s was already pretty much confined to modernday Guiné Bissau and Casamance (a region in southern Senegal which only was ceded to the French in 1888 and where a Portuguese-based Creole is still being spoken!).

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Cape Verde Slave Census of 1856 (part 1)

40-anos-da-Independância-de-Cabo-Verde-Unilab-550x275

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Cape Verde, an independent country since July 5th 1975!
Cape Verdeans: an indomitable people for more than 500 years!

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Cape Verde Slave Census of 1856

SUMMARY

Number of slaves 5,182
Creole (i.e. born in Cape Verde) 4,266 (82% of total)
African (mainland) 867 (17% of total)
African specified ethnically 130 (2,5% of total)

TOP 3 BREAKDOWN OF AFRICAN BORN SLAVES

Mandinga (Upper Guinea) 34 – 26% of African specified
Fula (Upper Guinea) 19 – 15% of African specified
Bijago (Guiné Bissau) 18 – 14% of African specified
Source: Carreira (1972)

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